largest lens opening (smallest f-number) at which a lens can be set. A fast lens transmits more light and has a larger opening than a slow lens. Determined by the maximum aperture of the lens in relation to its focal length. The speed of a lens is relative to it's focal length. A 400 mm lens with a maximum aperture of f/3.5 is considered extremely fast, while a 28mm f/3.5 lens is thought to be relatively slow.